| 15.64493 % | Casdin Capital, LLC | 7,207,165 | -500,000 | |
| 11.03182 % | BlackRock Institutional Trust Company, N.A. | 5,082,040 | 12,434 | |
| 9.21852 % | T. Rowe Price Investment Management, Inc. | 4,246,706 | 2,898,698 | |
| 7.50321 % | Invesco Advisers, Inc. | 3,456,512 | 126,618 | |
| 6.27128 % | The Vanguard Group, Inc. | 2,888,997 | 93,130 | |
| 3.05848 % | State Street Global Advisors (US) | 1,408,955 | -24,841 | |
| 2.70915 % | Fred Alger Management, LLC | 1,248,028 | -285,094 | |
| 2.39236 % | MFS Investment Management | 1,102,091 | 337,420 | |
| 2.22809 % | Rockefeller Capital Management | 1,026,415 | 62,158 | |
| 2.16553 % | Columbia Wanger Asset Management, LLC | 997,599 | 116,257 | |
| 2.13430 % | Dimensional Fund Advisors, L.P. | 983,208 | 62,935 | |
| 1.99949 % | Geode Capital Management, L.L.C. | 921,105 | 33,849 | |
| 1.99591 % | Palisade Capital Management, LLC | 919,459 | -27,626 | |
| 1.95750 % | Geneva Capital Management LLC | 901,764 | 49,637 | |
| 1.56316 % | Voya Investment Management LLC | 720,103 | 135,599 | |
| 1.52854 % | Cramer Rosenthal McGlynn, LLC | 704,155 | 19,770 | |
| 1.42916 % | RBC Global Asset Management (U.S.) Inc. | 658,371 | 47,947 | |
| 1.39622 % | Granahan Investment Management, LLC | 643,200 | 203,700 | |
| 1.38575 % | Mutual of America Capital Management LLC | 638,373 | 115,963 | |
| 1.17198 % | Gagnon Securities LLC | 539,896 | -1,056 | |
| 1.15423 % | Stephens Investment Management Group, LLC | 531,719 | 34,488 | |
| 1.13271 % | Royce Investment Partners | 521,805 | -338,939 | |
| 1.08415 % | Eagle Asset Management, Inc. | 499,435 | -101,716 | |
| 0.97327 % | Curi RMB Capital, LLC | 448,358 | 9,673 | |
| 0.91223 % | Citadel Advisors LLC | 420,237 | 297,435 | |
| 0.80462 % | Peregrine Capital Management, LLC | 370,663 | 6,217 | |
| 0.77696 % | Foord Asset Management (Singapore) Pte. Ltd. | 357,921 | 0 | |
| 0.76968 % | Northern Trust Investments, Inc. | 354,571 | 7,074 | |
| 0.75321 % | Charles Schwab Investment Management, Inc. | 346,982 | 6,156 | |
| 0.74996 % | Managed Account Advisors LLC | 345,487 | 34,089 | |
| 0.73119 % | Ranger Investment Management, L.P. | 336,840 | -770 | |
| 0.70819 % | BofA Global Research (US) | 326,244 | 63,265 | |
| 0.69371 % | Morgan Stanley & Co. LLC | 319,573 | 106,638 | |
| 0.66447 % | Renaissance Technologies LLC | 306,100 | 259,100 | |
| 0.64822 % | New York State Common Retirement Fund | 298,618 | 1,170 | |
| 0.62898 % | Carne Global Fund Managers (Ireland) Limited | 289,751 | 0 | |
| 0.59526 % | Gagnon Advisors, LLC | 274,217 | 0 | |
| 0.55373 % | Mathew (Aby J) | 255,087 | -536 | |
| 0.53937 % | Qube Research & Technologies Ltd | 248,474 | 238,533 | |
| 0.53876 % | Envestnet Asset Management, Inc. | 248,193 | -25,136 | |
| 0.49665 % | Next Century Growth Investors, LLC | 228,790 | -80,896 | |
| 0.49204 % | Uniplan Investment Counsel, Inc. | 226,668 | -20,710 | |
| 0.45580 % | JPMorgan Asset Management U.K. Limited | 209,973 | -11,555 | |
| 0.42003 % | Russell Investments Trust Company | 193,497 | -64,270 | |
| 0.39963 % | Principal Global Investors (Equity) | 184,099 | -75,389 | |
| 0.36937 % | Parametric Portfolio Associates LLC | 170,159 | -2,144 | |
| 0.35796 % | BlackRock Advisors (UK) Limited | 164,901 | 29 | |
| 0.34025 % | BlackRock Asset Management Ireland Limited | 156,745 | 47,140 | |
| 0.33316 % | Bard Financial Services, Inc. | 153,475 | 0 | |
| 0.32966 % | Foster (Karen A) | 151,865 | -400 | |
| 0.32927 % | Meros Investment Management, LP | 151,687 | 20,037 | |
| 0.31992 % | Raymond James & Associates, Inc. | 147,380 | -86,708 | |
| 0.31322 % | TD Asset Management Inc. | 144,290 | 105,470 | |
| 0.29414 % | UBS Switzerland AG | 135,500 | 73,391 | |
| 0.29027 % | UBS Financial Services, Inc. | 133,721 | -20,724 | |
| 0.28681 % | Isthmus Partners, LLC | 132,125 | 509 | |
| 0.28440 % | BlackRock Investment Management, LLC | 131,013 | 24,685 | |
| 0.27889 % | Mellon Investments Corporation | 128,477 | -2,004 | |
| 0.27452 % | PSP Investments | 126,465 | 11,769 | |
| 0.25792 % | de Greef (Roderick) | 118,816 | -112,835 | |
| 0.25118 % | D. E. Shaw & Co., L.P. | 115,712 | 115,712 | |
| 0.24914 % | Richardson (Garrie) | 114,773 | -3,070 | |
| 0.24511 % | SEI Investments Management Corporation | 112,913 | -329 | |
| 0.24303 % | Goldman Sachs Asset Management, L.P. | 111,958 | -176,253 | |
| 0.22922 % | Legal & General Investment Management Ltd. | 105,595 | 10,001 | |
| 0.22506 % | Macquarie Investment Management | 103,677 | -1,262 | |
| 0.22400 % | BlackRock Financial Management, Inc. | 103,192 | -8,309 | |
| 0.22095 % | Wichterman (Troy) | 101,784 | -877 | |
| 0.21857 % | State of Wisconsin Investment Board | 100,688 | 15,324 | |
| 0.21381 % | Berard (Todd) | 98,494 | -304 | |
| 0.21025 % | Invesco Capital Management LLC | 96,855 | -97,461 | |
| 0.20177 % | Wells Fargo Advisors | 92,950 | 6,869 | |
| 0.20168 % | Rhumbline Advisers Ltd. Partnership | 92,908 | 9,681 | |
| 0.19822 % | Barings LLC | 91,316 | 0 | |
| 0.18898 % | Jane Street Capital, L.L.C. | 87,056 | 39,275 | |
| 0.18107 % | Allspring Global Investments, LLC | 83,415 | -10,604 | |
| 0.17968 % | Columbia Threadneedle Investments (US) | 82,772 | -6,298 | |
| 0.16733 % | Rathbones Investment Management Limited | 77,086 | 0 | |
| 0.16651 % | BNY Mellon Asset Management | 76,707 | 4,086 | |
| 0.16064 % | Barclays Bank PLC | 74,002 | 26,245 | |
| 0.15608 % | Schweizerische Nationalbank | 71,900 | 0 | |
| 0.14937 % | Squarepoint Capital LLP | 68,809 | 68,809 | |
| 0.14648 % | UBS Asset Management (Switzerland) | 67,479 | 29,369 | |
| 0.14522 % | RBC Dominion Securities, Inc. | 66,899 | 66,899 | |
| 0.14379 % | Nuveen LLC | 66,241 | -13,317 | |
| 0.14340 % | Goldman Sachs Asset Management International | 66,060 | 66,060 | |
| 0.14114 % | Conestoga Capital Advisors, LLC | 65,017 | 0 | |
| 0.14106 % | Zacks Investment Management, Inc. | 64,984 | 63,226 | |
| 0.13813 % | RBC Capital Markets Wealth Management | 63,634 | -21,645 | |
| 0.13055 % | Tudor Investment Corporation | 60,139 | -265 | |
| 0.12677 % | Aquatic Capital Management LLC | 58,397 | 58,397 | |
| 0.12497 % | New York State Teachers' Retirement System | 57,569 | 0 | |
| 0.12019 % | ClariVest Asset Management LLC | 55,368 | -15 | |
| 0.11433 % | LPL Financial LLC | 52,667 | 4,834 | |
| 0.11192 % | Aebersold (Sarah) | 51,560 | -198 | |
| 0.10586 % | Citi Investment Research (US) | 48,767 | -8,751 | |
| 0.10337 % | Cerity Partners LLC | 47,621 | -982 | |
| 0.10084 % | Susquehanna International Group, LLP | 46,454 | -2,402 | |
| 0.09621 % | Dimensional Fund Advisors, Ltd. | 44,319 | 927 | |
| 0.09562 % | Aperio Group, LLC | 44,049 | -14,288 | |