| 17.66521 % | Liontrust Investment Partners LLP | 11,615,620 | -17,832 | |
| 16.08618 % | Aberforth Partners LLP | 10,577,337 | 0 | |
| 10.24571 % | Robert Walters Employee Benefit Trust | 6,736,987 | 0 | |
| 6.92487 % | Wellcome Trust | 4,553,390 | 918,800 | |
| 6.18489 % | Premier Asset Management Ltd | 4,066,824 | 2,766,824 | |
| 6.08237 % | Schroder Investment Management Ltd. (SIM) | 3,999,412 | 347,578 | |
| 6.00519 % | abrdn Investment Management Limited | 3,948,664 | 0 | |
| 5.31063 % | Fidelity International | 3,491,962 | -135,065 | |
| 4.73845 % | Aegon Asset Management Ltd | 3,115,729 | 0 | |
| 4.44162 % | Canaccord Genuity Wealth Management | 2,920,552 | 0 | |
| 3.85375 % | Jupiter Asset Management Ltd. | 2,534,004 | 0 | |
| 3.78762 % | INVESCO Asset Management Limited | 2,490,517 | 0 | |
| 3.12596 % | Walters (Robert Charles) | 2,055,449 | 0 | |
| 2.17248 % | FIL Pensions Management | 1,428,500 | 294,130 | |
| 1.83106 % | Otus Capital Management Limited | 1,203,996 | 50,806 | |
| 1.74026 % | Standard Life Assurance Limited | 1,144,293 | -17,511 | |
| 1.02909 % | Bannatyne (Alan Robert) | 676,667 | 0 | |
| 0.95195 % | Dimensional Fund Advisors, L.P. | 625,946 | -42,096 | |
| 0.91539 % | Cape Wrath Capital Limited | 601,909 | 0 | |
| 0.91249 % | Martin Currie Investment Management Ltd. | 600,000 | -200,000 | |
| 0.76041 % | Chelverton Asset Management Ltd. | 500,000 | 0 | |
| 0.71052 % | Legal & General Investment Management Ltd. | 467,197 | 0 | |
| 0.69136 % | Artemis Investment Management LLP | 454,600 | -141,536 | |
| 0.57703 % | Vanguard Global Advisers LLC | 379,424 | 21,458 | |
| 0.54380 % | AXA Investment Managers UK Ltd. | 357,573 | -57,175 | |
| 0.47285 % | M&G Investment Management Ltd. | 310,919 | -67,400 | |
| 0.41755 % | Martin (Mark) | 274,554 | -65,047 | |
| 0.37240 % | Dimensional Fund Advisors, Ltd. | 244,866 | 992 | |
| 0.33921 % | Koska Investments Limited | 223,042 | 0 | |
| 0.21385 % | abrdn Alternative Investments Limited | 140,615 | 0 | |
| 0.21385 % | BlackRock Advisors (UK) Limited | 140,615 | 0 | |
| 0.18668 % | The National Farmers Union Mutual Insurance Society Limited | 122,750 | -3,270 | |
| 0.18460 % | Norges Bank Investment Management (NBIM) | 121,383 | -3,206 | |
| 0.15710 % | TrinityBridge | 103,300 | -36,800 | |
| 0.15208 % | Teviot Partners LLP | 100,000 | 0 | |
| 0.14448 % | JPMorgan Asset Management U.K. Limited | 95,000 | 12,000 | |
| 0.13687 % | Waystone Management Company (Lux) SA | 90,000 | 0 | |
| 0.13096 % | River Global Investors LLP | 86,110 | 19,500 | |
| 0.11862 % | Van de Walle (Leslie) | 78,000 | 18,500 | |
| 0.10316 % | abrdn Investments Limited | 67,832 | -650,071 | |
| 0.09060 % | Daubeney (Giles Philip) | 59,574 | 0 | |
| 0.08511 % | VM Vermögens-Management GmbH | 55,961 | 0 | |
| 0.08001 % | Gibson (Kevin Joseph) | 52,607 | 0 | |
| 0.07967 % | Scottish Equity Partners LLP | 52,389 | 450 | |
| 0.07604 % | Fidelity Institutional Asset Management | 50,000 | 0 | |
| 0.07038 % | State Street Global Advisors (UK) Ltd. | 46,275 | -11,608 | |
| 0.05037 % | Sims (Lindsay Caroline) | 33,121 | 0 | |
| 0.04562 % | Duncan (Celia Mary) | 30,000 | 0 | |
| 0.04562 % | Bower (David) | 30,000 | 0 | |
| 0.04562 % | Marsh (John Bernard) | 30,000 | 0 | |
| 0.04473 % | Abbas (Simon Robert) | 29,411 | 0 | |
| 0.03513 % | AFH Wealth Management | 23,099 | 859 | |
| 0.03205 % | Fieldsend (Jonathan) | 21,071 | 9,062 | |
| 0.03042 % | Lo (Nicholas Charles Timothy) | 20,000 | 0 | |
| 0.02668 % | Rogerson (Neil) | 17,542 | 0 | |
| 0.02541 % | Westlake (Dolly) | 16,707 | 4,979 | |
| 0.02451 % | Mitchison (Robert Elliott) | 16,116 | 1,168 | |
| 0.02302 % | Ross (Andrea) | 15,138 | 0 | |
| 0.02294 % | Buchanan (James Gordon Nicholas) | 15,085 | 0 | |
| 0.02281 % | Rothschild & Co. Bank AG | 15,000 | 0 | |
| 0.02281 % | Rachet (Coralie) | 15,000 | 0 | |
| 0.02256 % | Isenberg (A) | 14,831 | 0 | |
| 0.02121 % | Mlanao (Alain Amadi) | 13,945 | 0 | |
| 0.02059 % | Webber (Rebecca) | 13,540 | 0 | |
| 0.02029 % | Simmonite (Daniel) | 13,344 | 0 | |
| 0.01836 % | Benson (Terry) | 12,074 | 0 | |
| 0.01679 % | Columbia Threadneedle Investments (UK) | 11,039 | 0 | |
| 0.01621 % | DFA Australia Ltd. | 10,656 | -6,703 | |
| 0.01470 % | Ashley (Matthew) | 9,667 | 0 | |
| 0.01065 % | Hesmondhalgh (Jane) | 7,000 | 7,000 | |
| 0.00912 % | Dodge (Tanith Claire) | 6,000 | 0 | |
| 0.00175 % | Dimensional Fund Advisors Canada ULC | 1,149 | 0 | |
| 0.00002 % | Aviva Investors Global Services Limited | 14 | 0 | |
| 0 % | Daubeney (Sophie) | 0 | -17,500 | |
| 0 % | Daubeney (Isabel) | 0 | -17,500 | |
| 0 % | Tudor (Monica) | 0 | -15,257 | |
| 0 % | Thesis Asset Management Limited | 0 | -480,216 | |
| 0 % | Oddo BHF Asset Management S.A.S | 0 | -10,000 | |
| 0 % | Wesleyan Assurance Society | 0 | -50,000 | |
| 0 % | Acadian Asset Management LLC | 0 | -58,396 | |
| 0 % | HSBC Global Asset Management (UK) Limited | 0 | -61,675 | |
| 0 % | Royal London Asset Management Ltd. | 0 | -95,000 | |
| 0 % | Irish Life Investment Managers Ltd. | 0 | -11,246 | |
| 0 % | Invesco Advisers, Inc. | 0 | -128,937 | |
| 0 % | Victory Capital Management Inc. | 0 | -2,983 | |
| 0 % | Mellon Investments Corporation | 0 | -118,365 | |
| 0 % | Dowgate Wealth Limited | 0 | -20,000 | |