| 7.81929 % | Portolan Capital Management, L.L.C. | 3,633,622 | 670,225 | |
| 6.72980 % | Columbia Threadneedle Investments (UK) | 3,127,338 | 128,326 | |
| 5.69809 % | BlackRock Institutional Trust Company, N.A. | 2,647,901 | -36,107 | |
| 5.44211 % | The Vanguard Group, Inc. | 2,528,950 | 3,271 | |
| 4.49645 % | Kent Lake PR LLC | 2,089,500 | -211,500 | |
| 4.01171 % | Walt (David R) | 1,864,240 | 2,594 | |
| 2.46375 % | Invenomic Capital Management LP | 1,144,904 | 242,861 | |
| 2.42733 % | William Blair Investment Management, LLC | 1,127,981 | 21,918 | |
| 2.38765 % | Columbia Threadneedle Investments (US) | 1,109,539 | 393,313 | |
| 2.33190 % | Easterly Investment Partners LLC | 1,083,635 | -241,985 | |
| 2.06100 % | Blue Water Life Science Advisors, LP. | 957,747 | 585,572 | |
| 1.90893 % | Geode Capital Management, L.L.C. | 887,080 | 21,210 | |
| 1.72958 % | Dimensional Fund Advisors, L.P. | 803,735 | -37,598 | |
| 1.65225 % | Balyasny Asset Management LP | 767,801 | 723,295 | |
| 1.64591 % | State Street Investment Management (US) | 764,855 | 3,547 | |
| 1.48181 % | Trium Capital LLP | 688,597 | 102,172 | |
| 1.45473 % | Williams Jones Wealth Management LLC | 676,014 | 162,525 | |
| 1.35944 % | Morgan Stanley & Co. LLC | 631,732 | 180,440 | |
| 1.23008 % | Tikvah Management LLC | 571,619 | 0 | |
| 1.11038 % | Parkman Healthcare Partners LLC | 515,992 | 515,992 | |
| 0.95587 % | Marshall Wace LLP | 444,191 | 444,191 | |
| 0.90565 % | abrdn Inc. | 420,856 | 152,139 | |
| 0.82839 % | UBS Financial Services, Inc. | 384,951 | -730,027 | |
| 0.81587 % | Two Sigma Investments, LP | 379,137 | 229,674 | |
| 0.76251 % | SCS Capital Management LLC | 354,339 | 0 | |
| 0.70083 % | Schonfeld Strategic Advisors LLC | 325,677 | 325,677 | |
| 0.65802 % | Northern Trust Investments, Inc. | 305,784 | -2,754 | |
| 0.64955 % | Meister Paul M | 301,844 | 2,242 | |
| 0.64340 % | Charles Schwab Investment Management, Inc. | 298,989 | 6,184 | |
| 0.62354 % | State of Wisconsin Investment Board | 289,759 | 161,751 | |
| 0.61839 % | Toloue (Masoud) | 287,366 | -1,242 | |
| 0.60378 % | Citi Investment Research (US) | 280,578 | 254,082 | |
| 0.52314 % | J.P. Morgan Securities LLC | 243,101 | -27,609 | |
| 0.49839 % | Envestnet Asset Management, Inc. | 231,601 | -268,797 | |
| 0.33132 % | Goldman Sachs & Company, Inc. | 153,964 | -107,760 | |
| 0.32544 % | RBC Capital Markets Wealth Management | 151,230 | 2,261 | |
| 0.30119 % | Jane Street Capital, L.L.C. | 139,964 | 81,651 | |
| 0.29740 % | BlackRock Financial Management, Inc. | 138,200 | -78,230 | |
| 0.29726 % | Donnelly (William P) | 138,137 | 5,076 | |
| 0.29242 % | Parametric Portfolio Associates LLC | 135,888 | 112,032 | |
| 0.27252 % | Jump Financial, LLC | 126,641 | 21,184 | |
| 0.27250 % | Fidelity Management & Research Company LLC | 126,632 | 28,692 | |
| 0.27099 % | BlackRock Investment Management, LLC | 125,931 | 21,577 | |
| 0.26680 % | Empire Financial Management Company, LLC | 123,980 | 73,980 | |
| 0.25633 % | Wellington Management Company, LLP | 119,115 | 119,115 | |
| 0.25402 % | Cubist Systematic Strategies, LLC | 118,043 | -169,188 | |
| 0.24771 % | Wells Fargo Advisors | 115,113 | 91,688 | |
| 0.22208 % | Renaissance Technologies LLC | 103,200 | -99,721 | |
| 0.21519 % | ACATIS Investment Kapitalverwaltungsgesellschaft GmbH | 100,000 | 0 | |
| 0.21488 % | Goldman Sachs Asset Management, L.P. | 99,856 | 11,884 | |
| 0.18993 % | AQR Capital Management, LLC | 88,259 | -10,731 | |
| 0.18553 % | American Century Investment Management, Inc. | 86,215 | 7,085 | |
| 0.16842 % | Barclays Bank PLC | 78,266 | 24,818 | |
| 0.15393 % | Dimensional Fund Advisors, Ltd. | 71,529 | -4,725 | |
| 0.15291 % | Nuveen LLC | 71,058 | -342,523 | |
| 0.14827 % | BNY Mellon Asset Management | 68,902 | 3,080 | |
| 0.13988 % | Graypoint LLC | 65,000 | 0 | |
| 0.13984 % | Invesco Capital Management LLC | 64,984 | 21,167 | |
| 0.13842 % | Susquehanna International Group, LLP | 64,326 | 8,059 | |
| 0.13355 % | The Clarius Group, LLC | 62,060 | 0 | |
| 0.13299 % | BofA Global Research (US) | 61,801 | -75,594 | |
| 0.13047 % | Rhumbline Advisers Ltd. Partnership | 60,629 | 4,094 | |
| 0.12912 % | Mount Lucas Management LP | 60,000 | 16,163 | |
| 0.12056 % | XTX Markets LLC | 56,024 | 33,007 | |
| 0.11327 % | Hlavinka (Sarah E) | 52,635 | 2,615 | |
| 0.11213 % | Peddock Capital Advisors, LLC | 52,105 | 34,503 | |
| 0.10258 % | PDT Partners, LLC | 47,668 | 47,668 | |
| 0.09572 % | AllianceBernstein L.P. | 44,480 | 7,500 | |
| 0.09386 % | RBC Wealth Management, International | 43,616 | 21,840 | |
| 0.09030 % | Quantinno Capital Management LP | 41,964 | 31,664 | |
| 0.08570 % | Avantis Investors_NLE | 39,827 | 37,990 | |
| 0.08549 % | Mellon Investments Corporation | 39,726 | 1,802 | |
| 0.08521 % | T. Rowe Price International Ltd | 39,599 | 1,665 | |
| 0.07532 % | Credit Industriel et Commercial SA | 35,000 | 0 | |
| 0.07405 % | BNP Paribas Securities Corp. North America | 34,410 | -152,476 | |
| 0.06857 % | Quantbot Technologies, LP | 31,864 | 20,592 | |
| 0.06445 % | HSBC Global Asset Management (UK) Limited | 29,949 | -1,184 | |
| 0.06272 % | Nebula Research & Development LLC | 29,145 | -6,820 | |
| 0.05682 % | Flynn (Karen A) | 26,405 | 0 | |
| 0.05234 % | DWS Investments UK Limited | 24,321 | -6,394 | |
| 0.05128 % | MetLife Investment Management, LLC | 23,830 | -153 | |
| 0.05084 % | Wells Fargo Bank, N.A. | 23,624 | 2,669 | |
| 0.04583 % | Ironwood Investment Management, LLC | 21,297 | 21,297 | |
| 0.04411 % | Osaic Holdings, Inc. | 20,499 | 1,396 | |
| 0.04399 % | Mirae Asset Global Investments (USA) LLC | 20,444 | -936 | |
| 0.04354 % | Magovcevic-Liebisch (Ivana) | 20,235 | 7,435 | |
| 0.04304 % | Ersel Asset Management SGR S.p.A. | 20,000 | 0 | |
| 0.03879 % | T. Rowe Price Associates, Inc. | 18,025 | 5,300 | |
| 0.03788 % | Corebridge Financial Inc. | 17,602 | -262 | |
| 0.03758 % | Commonwealth Financial Network | 17,465 | -450 | |
| 0.03744 % | JPMorgan Private Bank (United States) | 17,397 | 405 | |
| 0.03635 % | INTECH Investment Management LLC | 16,893 | -6,585 | |
| 0.03633 % | Prelude Capital Management, LLC | 16,884 | -4,126 | |
| 0.03381 % | Tallon Kerry Patrick | 15,710 | 15,710 | |
| 0.03343 % | Metropolitan Life Insurance Co. (US) | 15,533 | 0 | |
| 0.03308 % | Squarepoint Capital LLP | 15,372 | -1,150 | |
| 0.03243 % | Sriram (Vandana) | 15,072 | -229 | |
| 0.03188 % | State of Wyoming | 14,814 | 7,599 | |
| 0.03121 % | Brevan Howard Capital Management LP | 14,505 | 14,505 | |
| 0.03110 % | Moors & Cabot Inc. | 14,452 | -18,001 | |